2
I?IINOA?NOAI IOI?IIE CAeI?IA’ss OE?A?IE
OA?IIIIEUeNEA AeA?AEAAIA IAAeE*IA
AEAAeAIIss II. I.ss.AI?AA*AANUeEIAI
UA?AAEIA IAENEI NOAIAIIAE*
OAeE 611.36:57.017 645-08:616-097] 591.4
INIAEEAINOI AOAeIAE IA*IIEE UO?IA
A ?AIIUeIIO IINOIAOAEUeIIIO IA?IIAeI
A II?II OA IINEss AIOO?IOIUeI*A?AAIIAI AAAAeAIIss AIOEAAIIA
(aiaoiii-aenia?eiaioaeueia aeineiaeaeaiiy)
14.03.01. – ii?iaeueia aiaoiiiy
AAOI?AOA?AO
aeena?oaoei? ia caeiaoooy iaoeiaiai nooiaiy
eaiaeeaeaoa iaaee/ieo iaoe
Oa?iiiieue – 1999
Aeena?oaoei?th ? ?oeiien. ?iaioa aeeiiaia a Caii?icueeiio aea?aeaaiiio
iaaee/iiio oiiaa?neoaoi IIC Oe?a?ie ia eaoaae?i ii?iaeueii? aiaoiii?.
Iaoeiaee ea?iaiee: aeieoi? iaaee/ieo iaoe, i?ioani?
Aieioei Ieeiea Aiaoieieiae/
caaiaeoaa/ eaoaae?e ii?iaeueii? aiaoiii?
Caii??cueeiai aea?aeaaiiai
iaaee/iiai oi?aa?neoaoo
Io?oe?ei? iiiiaioe:
aeieoi? iaaee/ieo iaoe, i?ioani?
Eoie? Aieoi? Ieoaeeiae/ – caa?aeoaa/ eaoaae?e ii?iaeueii? aiaoii?.
Oa?eianueeiai aea?aeaaiiai iaaee/iiai oiiaa?neoaoo
aeieoi? iaaee/ieo iaoe, i?ioani?
Aieiaaoeueeee Aiae?ie Noaiaiiae/ – , caaiaeoaa/ eaoaae?e, aiaoiii?
ethaeeie, iia?aoeaii? oi?o?ai? oa ainoieiai?. Oaeai?iaenueeiai
aea?aeaaiiai oi?aa?neoaoo
I?iaiaeia onoaiiaa:
Eoaainueeee aea?aeaaiee iaaee/iee oiiaa?neoao IIC OE?a?ie, eaoaae?a
ii?iaeueii? aiaoiii?, i. Eoaainuee.
Caoeno aiaeaoaeaoueny ” 20 ” o?aaiy 1999 ?. i ” 14-00 ” aiaeeii
ia caniaeaiii niaoeiaeiciaaii? a/aii? ?aaee E 58.601.01
o Oa?iiiieuenueeie aea?aeaaiie iaaee/iie aeaaeaii? ii. I.Ai?aa/aanueeiai
(i. Oa?iiiieue, iaeaeai Aiei,1)
C aeena?oaoei?th iiaeia iciaeiieoenue o aiaeiioaoei Oa?iiiieuenueei?
aea?aeaaii? iaaee/ii? aeaaeaii? ii. I.Ai?aa/aanueeiai
(i. Oa?iiiieue, aoe. ?onueea, 12)
Aaoi?aoa?ao ?icineaiee ” 20 ” ea?oiy 1999 ?.
A/aiee nae?aoa?
niaoeiaeiciaaii? a/aii? ?aaee
aeieoi? iaaee/ieo iaoe, i?ioani?
Eio I.I
CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE
Aeooaeueiinoue i?iaeaie. O ca’yceo c ?icaeoeii iaoee oa oaoiiee,
eaaeiiiiaeiaiith iiyaith iiaeo aiieiai/ieo oa neioaoe/ieo iieiia?ia, ui
ciaoiaeyoue oe?iea canoinoaaiiy ye o i?iieneiainoi, oae i a ia?iaeiiio
ainiiaea?noai, niinoa?iaa?oueny iiai?oaiiy aeieiai/iiai noaio ia
ieaiaoi, i, ye ianeiaeie, caieueoaiiy iaaaioaaeaiiy ia iioio nenoaio ye
ethaeeie, oae i oaa?ei (E?eooioi?iaa A.A.,1998). Ana oea iaaaoeaii
aieeaa? ia noaiiaeaiiy i?aaiicio a oeieiio o i?aiaoaeueiiio oa ?aiiueio
iinoiaoaeueiiio ia?iiaei ?icaeoeo (Aantheiaa E.E.,1997) oa ia eiai
iioiio nenoaio.
Ia/iiea, yea ? aeaea?aeii iieiiioaioieo noiaao?iaeo eeioei, o
i?aiaoaeueiiio ia?iiaei ?icaeoeo aeeiio? ?ieue o?aiceoi?iiai
oeaio?aeueiiai eiioi?aeiiai i?aaio, (Aeaenaaaa E.I., A?ucaeia O.I.,
Iaaeiae/ N.E.,Eeue/aae/ I.A., 1991) iicaoaath/enue oei?? ooieoei? i?e
ia?aoiaei aei iica/a?aaiiai inioaaiiy, ui noi?iaiaeaeo?oueny oi?ioaaiiyi
?? ye aeaoieneeaoeieiiai oa aecie?eiiiai i?aaio. Ii?oiooieoe?iee noai
ia/iiee i?iiiiothoue ?icaeyaeaoe ye ia?ea? aeieiai/iiai i?aneiao ia
i?aaiici a oeieiio (Ai?iaeei Ae.E., Ie/o?eia N.A.,1998).
A eioa?aoo?i a?aeo? aeineiaeaeaiue i?i aeeiaiieo eiioi?aeiiai eiiiiiaioo
ia/iiee oa iaoaiicie i?eieiaiiy a i?aaii yaeu eiioioeeoiiiaco.
Ia?aaaaeii aea/aee ciiie ii?ei?aeiiai ?inoea ye ?aaeoeith ia oiene/io
aeith ?icieo oaeoi?ia (Auaiei I.E., Ia?ouiaiei E.A., 1988, Ieeiiaiei
A.A., 1994, Ohneda O., Bautch V.L., 1997). Cie?aia, iaia? aiaeiiinoae
i?i aieea ia noai ia/iiee, ye o?aiceoi?iiai i?aaio iioiioaoo,
aioo?ioiuei/a?aaii? aioaaiii? ae??, yea aeeeeea? i?enei?aiiy oi?ioaaiiy
ii?oiooieoeiiiaeueieo cii eiioi?aeieo i?aaiia (Aieioei I.A., 1991,1995,
Nu?oeia A.E., 1991, Ea?cia I.A., 1994).
Anoaiiaeaiiy caeiiiii?iinoae ii?oiaaiaco eiioi?aeii? oeaieie ia/iiee a
ii?ii oa iiney aioo?ioiuei/a?aaiiai aaaaeaiiy aioeaaiia aeicaieeoue
iiaeeaeoe oyaeaiiy i?i ia/iieo ye eiioi?aeiee i?aai oa oaeineiiaeeoe
iaoiaee ae?aaiinoeee aioaiaoaeueii? aei? /oaea?iaeieo aioeaaiia oa
iiaeieoe aei ?ice?eooy iaoaiiciia oi?ioaaiiy iioiieiai/ii? oiea?aioiinoi
o iiaiia?iaeaeaieo.
Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie. ?iaioa aeeiiaia
o ?aieao iaoeiai-aeineiaeii? ?iaioe eaoaae?e ii?iaeueii? aiaoiii?
Caii?icueeiai aea?aeaaiiai iaaee/iiai oiiaa?neoaoo “Caeiiiii?iinoi
ii?oiaaiaco eiioi?aeeo i?aaiia a ii?ii oa iiney aioo?ioiuei/a?aaii?
aioeaaiii? noeioeyoei? i caniae ei?aeoei? iioiieiai/iiai aeaoioeeoo ca
aeiiiiiaith iioiiiiaeoeyoi?ia” (N aea?ae?a?no?aoei? 0194 U 008429).
Iaoa aeineiaeaeaiiy. Aea/eoe caeiiiii?iinoi ii?oiaaiaco ia/iiee
iiaiia?iaeaeaieo uo?ia oa aoaeiae ?? eiioi?aeiiai eiiiiiaioo a ii?ii oa
iiney aioo?ioiuei/a?aaiiiai aaaaeaiiy aioeaaiia.
Aeey aeinyaiaiiy iaoe aeineiaeaeaiiy aoee aecia/ai? ianooiii caaaeaiiy:
1. Aea/eoe aeeiaiieo aaniethoii? oa aiaeiinii? iane ia/iiee
iiaiia?iaeaeaieo uo?ia a ii?ii oa iiney aioo?ioiuei/a?aaiiai aaaaeaiiy
aioeaaiia.
2. Anoaiiaeoe eieaeicaoeith oa i?eioeei aoaeiae eiioi?aeiiai eiiiiiaioo
ia/iiee iiaiia?iaeaeaieo uo?ia o ii?ii oa eiai ?aaeoeith ia
aioo?ioiuei/a?aaia aaaaeaiiy aioeaaiia.
3. Aecia/eoe iniaeeainoi noaiiaeaiiy ia/iieiaeo /anoi/ie
iiaiia?iaeaeaieo uo?ia iiney aioo?ioiuei/a?aaiiai aaaaeaiiy aioeaaiia.
4. Aea/eoe i?ieioa?aoeaio aeoeaiinoue aaiaoioeeoia uo?ia o ?aiiueiio
iinoiaoaeueiiio ia?iiaei a ii?ii oa iiney aei? aioeaaiia, ui aoee
oaaaeaii aioo?ioiuei/a?aaii.
5. Aeineiaeeoe ?iciiaeie aeieii?ioa?aeia o aaiaoioeeoao iioaeoieo oa
aenia?eiaioaeueieo uo?ia.
6. Anoaiiaeoe iniaeeainoi aeeiaiiee eeioei ic ?aoeaioi?aie aei eaeoeio
a?aoino a ia/iioei iioaeoieo oaa?ei oa uo?ia, yei caciaee
aioo?ioiuei/a?aaiiai aaaaeaiiy aioeaaiia.
Iaoeiaa iiaecia iaea?aeaieo ?acoeueoaoia. A ?iaioi aia?oa iienaii
caeiiiii?iinoi oiiia?aoi? oa aeeiaiiee eiioi?aeii? oeaieie ia/iiee a
?aiiueiio iineyiaoaeueiiio ia?iiaei o iioaeoieo oa aenia?eiaioaeueieo
uo?ia. Aia?oa anoaiiaeaii ii?oieiaith oa oiiia?aoith eiioi?aeiiai
eiiiiiaioo ia/iiee o iiaiia?iaeaeaieo, yeee i?aaenoaaeaiee ae?iaieie,
na?aaeiiie oa aaeeeeie eiioi?aeieie neoi/aiiyie oa iaeeie/ieie aeeoocii
?icoaoiaaieie eiioioeeoaie. O iioaeoieo uo?ia niinoa?iaa?oueny iinooiiaa
ciaioaiiy ieiui, caeiaii? eiioi?aeieie neoi/aiiyie, aei 14-? aeiae
aeeooy. Aioo?ioiuei/a?aaia aaaaeaiiy aioeaaiia, iacaeaaeii aiae ?o
i?e?iaee, i?enei?th? no?iee aiia?aoei? eiioioeeoia ic i?aaio oa
ciaioaiiy ieiui eiioi?aeiiai eiiiiiaioo.
Anoaiiaeaiee aieea aioeaaiii? aei? ia no?iee oi?ioaaiiy
no?oeoo?ii-ooieoeiiiaeueieo iaeeieoeue ia/iiee, ui i?iyaey?oueny o
iiaeaeuaiii i?ieioa?aoeaii? aeoeaiinoi aaiaoioeeoia o iiaiia?iaeaeaieo,
i?enei?aiii oii?yaeeoaaiiy oi?ioaaiiy ia/iieiaeo aaeie oa
noi?iaiaeaeo?oueny caieueoaiiyi aaniethoii? oa aiaeiinii? iane ia/iiee a
?aiiueiio iineyiaoaeueiiio ia?iiaei.
Aia?oa aeiaaaeaii caieueoaiiy aia?aaoe/ieo aeo?ao i?aaiicio o
aiaeiiaiaeue ia aioo?ioiuei/a?aaia aaaaeaiiy aioeaaiia, ui i?iyaey?oueny
o ciaioaiii aiinoo aeieiaaio a oeeoiieacii aaiaoioeeoia c iiaeaeueoith
eiiiainaoi?iith aeoeaaoei?th aeieiaaineioaoe/ii? ooieoei?. Aia?oa
iienaii no?oeoo?e ia/iiee, yei iathoue ?aoeaioi?e aei eaeoeio a?aoino –
aiaeioaeiioeeoe, eiioioeeoe, eeioeie Eoioa?a, oieaiaiioeeoe. Anoaiiaeai?
caeiiiii?iinoi ?o aeeiaiiee – iinooiiaa c?inoaiiy eieueeinoi
eaeoei-iiceoeaieo aiaeioaeiioeeoia oa ciaioaiiy aiinoo PNA+-eiioioeeoia
oa PNA+-eeioei Eoioa?a, ui aiaeia?aaea? i?ioeane noaiiaeaiiy
aeaoiiioeaii? no?oeoo?e i?aaio.
I?aeoe/ia cia/aiiy iaea?aeaieo ?acoeueoaoia. Iienaii iniaeeainoi
aeeiaiiee eiioi?aeiiai eiiiiiaioo ia/iiee o iiaiia?iaeaeaieo a ii?ii oa
iiney aioo?ioiuei/a?aaiiai aaaaeaiiy aioeaaiia iiaeooue neoaoaaoe
aeiaaiinoe/iei e?eoa?i?i aioo?ioiuei/a?aaii? aei? aioeaaiia ia i?aaiici
ieiaeo.
?acoeueoaoe uiaei ?icaeoeo o?aiceoi?ii? aaiaoiiaaaei? aianeiaeie
aioaiaoaeueiiai aaaaeaiiy aioeaai?a oa iniaeeainoi noaiiaeaiiy
ia/iieiaeo /anoi/ie oa iniiaieo ooieoeie i?aaio iiaeooue i?aaenoaaeyoe
iioa?an aeey ii?oieiaia, iioiieiaia, iaaeiao?ia i?e eieoaaiii aeioae c
a?iaeaeaieie iaoieiaiyie, oa iioaeoeiiiinoia i iioiieiaia i?e ?ic?iaoei
a?aoieia iioiicaoei? aeioae ?aiiueiai aieo, yei aioo?ioiuei/a?aaii
caciaee aieeao /oaea?iaeieo aioeaaiia.
Aeaii i?i ?iciiaeie no?oeoo?, yei iathoue ?aoeaioi?e aei eaeoeio
a?aoino a ia/iioei, iiaeooue aeei?enoiaoaaoeny a i?aeoeoei ?iaioe
iaoeiaeo eaai?aoi?ie iaaee/ieo oa iaoeiai-aeineiaeieo caeeaaeia.
?acoeueoaoe aeena?oaoei? ai?iaaaeaeaii a o/aiaee i?ioean ia eaoaae?ao
iiiaeueii? aiaoiii? Caii?iceiai aea?aeaaiiai iaaee/iiai oiiaa?neoaoo,
Caii?icueeiai iinoeoooo oaeineiiaeaiiy eiea?ia, Aoeiaeinueei? iaaee/ii?
aeaaeaii?, Oa?iiiieuenueei? iaaee/ii? aeaaeaii?, ?aaii-O?aieianueeiai
iaaee/iiai oiiaa?neoaoo, iaaee/iiai oaeoeueoaoo Oaeai?iaenueeiai
aea?aeaaiiai oiiaa?neoaoo.
Iniaenoee aianie caeiaoaa/a. Caeiaoaa/ai cae?eniaii ?ic?iaeo iniiaieo
oai?aoe/ieo ? i?aeoe/ieo iieiaeaiue ?iaioe. Aeanii?o/ii i?iaaaeaii
aenia?eiaio ii aioo?ioiuei/a?aaiiio aaaaeaiith aioeaaiia ieiaeai ia 18-o
aeiao i?aiaoaeueiiai ?icaeoeo, caaie aenia?eiaioaeueieo oaa?ei,
aeaeieaiiy iaoa?iaeo, aeaioiaeaiiy oa oa?aoaaiiy ainoieiai/ieo
i?aia?aoia ca icia/aieie a ?iaioi iaoiaeeeaie. Aaoi?ii aeeiiaii ia?aeiio
ia?iaeo aeaieo oa noaoenoe/io ia?iaeo ?acoeueoaoia, cae?eniaii
oioiaeieoiaioaoe?th, i?iaaaeaii aiae?c oa ocaaaeueiaiy ?acoeueoao?a,
noi?ioeueiaaii on? iieiaeaiiy ? aeniiaee ?iaioe.
Ai?iaaoeiy ?acoeueoaoia aeena?oaoei?. ?acoeueoaoe ?iaioe aoee
aeeeaaeai? oa iaaiai?ai? ia naiiia?i iieiaeeo a/aieo Caii?icueeiai
oiaa?enoaa AAA (1996, 1997), ia iiaeia?iaei?e iaoeiaie eiioa?aioei?
“Aeooaeueii ieoaiiy ii?oieiai?”, i?enay/aiie iai’yoi aeaaeaiiea Nii?uea
N.A., (Oa?iiiieue, 1996); ia ??-io iaoeiiiaeueiiio eiia?ani aiaoiiia,
ainoieiaia, aia?iieiaia oa oiiia?aoiaiaoiiia Oe?a?ie (Eoaainuee,1988);
ia iiaeia?iaeiie iaoeiai-ae?iaie/ie eiioa?aioei? “Ii?oieiaiy –
i?aeoe/iie aaoa?eia?i? oa iaaeeoeeii”, i?enay/aiie iai’yoi i?io.
Eiaaeuenueeiai I.I. (Aiea Oea?eaa, 1998); Versamlung der Anatomischen
Gesellschaft in Marburg (1994), 89.European Research Conference
(Blankenberge, Belgium, 1996); 15.Arbeitstagung der Anatomischen
Gesselshaft in Wurzburg (1998).
Ai?iaaoeiy aeena?oaoeieii? ?iaioe aoea i?iaaaeaia 5 ni/iy 1999 ?. ia
caniaeaiii Caii?icueeiai iaeaniiai iaoeiaiai oiaa?enoaa aiaoiiia,
ainoieiaia, aia?iieiaia oa oiiia?aoiaiaoiiia i. Caii??aeaey.
Ioaeieaoei?. Ca oaiith aeena?oaoei? iioaeieiaaii 12 ae?oeiaaieo ?iaio,
o?e c yeeo iaae?oeiaaii a oaoiaeo iaoeiaeo aeaeaiiyo AAE Oe?a?ie,
aeaa`youe – o ca??ieeao oa oacao eiioa?aioe?e, c’?cae?a.
Ianya oa no?oeoo?a aeena?oaoei?. Aeena?oaoeiy neeaaea?oueny ci anooio,
iaeyaeo eioa?aoo?e, ia?aeieo iaoa?iaeia oa iaoiaeia aeineiaeaeaiiy,
?icaeieo “?acoeueoaoe aeanieo aeineiaeaeaiue”, yeee neeaaea?oueny c
i’yoe iiae?icaeieia, iaaiai?aiiy io?eiaieo ?acoeueoaoia, aeniiaeia oa
nieneo aeei?enoaieo aeaea?ae eioa?aoo?e.
Aeena?oaoeieia ?iaioa aeeeaaeaia ia 172 noi?iieao iaoeiiieniiai oaenoo.
Iinoeoue 50 iethno?aoeie (40 iie?ioioia?aoie oa 10 aeiaa?ai), yei
caeiathoue 27 noi?iiie, oa 9 oaaeeoeue, yei caeiathoue 9 noi?iiie.
Ia?ae?e aeei?enoaieo eioa?aoo?ieo aeaea?ae iinoeoue 290 iaeiaioaaiue
(159 – e?a?i NIAe, 131 iioeo e?a?i) oa caeia? 23 noi?iiee iaoeiiieniiai
oaenoo.
INIIAIEE CI?NO ?IAIOE
Iaoa?iae oa iaoiaeeee aeineiaeaeaiiy. Ia’?eoaie aeineiaeaeaiiy noaea
ia/iiea 187 aieeo uo?ia eiii? Ainoa? 1-?, 3-?, 5-?, 7-?,11-?, 14-? oa
30-? aeiae iinoiaoaeueiiai aeeooy, eio?eo oo?eioaaee a oiiaao aiaa?ith.
Aoei aeineiaeaeaii 4 a?oie oaa?ei. Ia?oa – iioaeoia. Oaa?eiai ae?oai? oa
o?aouei? aenia?eiaioaeueieo a?oi ia 18-o aeiao aioo?ioiuei/a?aaiiai
ia?iiaeo ?icaeoeo aaiaeeee aioeaai. Uo?e /aoaa?oi?, eiio?ieueii?, a?oie
io?eioaaee oiciieiai/iee ?ic/ei o oi ae naii no?iee.
Iiaiia?iaeaeaii io?eiaii aiae naiie uo?ia ic aeaoiaaiei no?ieii
aaaioiinoi. Aioo?ioiuei/a?aaia aaaaeaiiy aioeaaiia oa oiciieiai/iiai
?ic/eio caeienithaaee iia?aoeaiei oeyoii ii caniao I.A.Aieioeia (1981).
Oaa?eiai ae?oai? a?oie aaiaeeee iioiiaeiaoeii ethaeeie ii?iaeueiee, ui
ia? aeia?i aioeaaiii aeanoeainoi c aeoaea iacia/iith oiene/iith,
ie?iaaiiith oa aae’thaaioiith aei?th. Aeey ii?iaiyiiy aieeao aioeaaiia
aieeiai? oa ai?onii? i?e?iaee ia ii?oiaaiac ia/iiee aeei?enoiaoaaee
aaeoeeio aeeao ia?ioeoio nooo, yeo aaiaeeee oaa?eiai o?aouei? a?oie a
aeici 25 AAAeA50 o 0,05 ie oiciieiai/iiai ?ic/eio oaa?eiai o?aouei?
a?oie. Iiaiia?iaeaeaii uo?yoa ia?iaeeeenue aeiiioaieie, a no?ie, ia
21-22 aeiao iiney ca/aooy.
Caaie oaa?ei i?iaiaeeee oeyoii aeaeaiioaoei? o ae?oaie iieiaeii aeiy, c
13.00 aei 14.00. Ia/iieo caaaeoaaee ia oi?ciiieo oa?acao oa ia/enethaaee
?? aaniethoio oa aiaeiinio iano, oienoaaee o noiioi Aoaia. Aiooaaee
na?ieii c?ice caaoiaoee 4-5 iei. Aeey iaeyaeiaiai ainoieiai/iiai e
ii?oiiao?e/iiai aeineiaeaeaiue aeei?enoiaoaaee oa?aoaaiiy aaiaoieneeiiii
oa aiciiii.
?iciiaeie aeieii?ioa?aeia aea/aee ca aeiiiiiaith O?E-?aaeoei? c
aeioa?aoaaiiyi yaea? aaiaoieneeiiii. Aiino aeieiaaio o oeeoiieacii
aaiaoioeeoia aecia/aee ca aeiiiiiaith oa?aoaaiiy ea?iiiii ca iaoiaeii
Aanoa. Iiae?aoiaoaaee eieueeinoue aaiaoioeeoia, a oeeoiieacii yeeo aoee
ae?iaii, na?aaeii oa aaeeei a?aioee aeieiaaio. Aeineiaeaeaiiy aiinoo
AeIE o yae?ao aaiaoioeeo?a i?iaiaeeee ca iaoiaeii Oueieueaaia c
iiaeaeueoith niaeo?ioioiiao?i?th ia aia?aoi “ETHIAI – E 2” i?e aeiaaeeii
oaeei 360 iie. No?oeoo?e, ui iinoyoue ?aoeaioi?e aei eaeoeio a?aoino
(PNA), aeyaeyee iioiiii?oieiai/iei iaoiaeii, aeei?enoiaoth/e
noaiaea?oiee iaai? IIE “Eaeoeioano” (i.Eueaia).
Ia c?icao ia/iiee iiae?aoiaoaaee aiaeiinio eieueeinoue aaiaoioeeoia ia
oiiaiie ieiui 3.06o103 iei2, aiino iiii- oa iieiioeeaa?ieo aaiaoioeeoia,
eeioei o noaii iioico, aiino iaia?aioiiaoicieo ee?oei (aiaeioaeiioeeoe,
ci?eiiiaeiaii ?aoeeoeiaiaeioaeiioeeoe, aiioaeiioeeoe aeia/ieo i?ioieia),
aiaeiinio ieiuo e?iaiiinieo noaeei ia/iiee, a oaeiae ieiuo, yeo caeiaee
ina?aaeee ii?eioeeoiiiaco oa eiioioeeoiiiaco. Na?aaeith ieiuo
no?oeoo?ii-ooieoeiiiaeueieo iaeeieoeue ia/iiee aecia/aee, aeoiaey/e c
aiaenoaii iiae vv.centralas, ui ? iaeaooiiie oeaio?aie ia/iieiaeo
/anoie.
Ii?oiiao?e/ii iieacieee aoei iaea?aeaii c aeei?enoaiiyi iaoiaeo
ii?oiiao?e/iiai eieueeiniiai iaeieo (Noaoaiia N.A., Eooi?aiei
I.N.,1988). Ani ?acoeueoaoe aeineiaeaeaiue ia?iaeyee iaoiaeaie
aa?iaoeieii? noaoenoeee. ?ica?aeiino? i?ae aeaiia na?aaei?ie aaaaeaee
a??ia?aeieie i?e ?? ???????????????
???????????yyy????.????1?????????????a
???&??yyyy?????AyyyOyyya?????????&?
?MathType??P????u th??????Aeyaeaiiy na?aae aaiaoioeeo?a
PNA+-eiioioeeoia, yei ? iioiieiai/ii iac?ieeie (Eooeee A.Ae.,1989), ? ua
iaeiei iiaeoaa?aeaeaiiyi inioaaiiy i?ioeania eiioioeeoiiiaco a ia/iioei
iiaiia?iaeaeaieo. Ciaioaiiy aiaeiinii? eieueeinoi eiioioeeoia a ia/?ioe?
iiaea aooe iia'ycaii ye c iia?aoei?th c i?aaio, oae i c iinooiiaei
aeic?iaaiiyi. Ineieueee ia oeueiio oiii ia niinoa?iaaeinue iiaeaeuaiiy
oaaioeeoa?ii? aeoeaiinoi oa ia c?inoaea eieueeinoue c?oeiiaaieo eeioei,
oi ciaioaiiy aiinoo eaeoei-iiceoeaieo eiioioeeoia o?aaa ?icaeyaeaoe ye
i?enei?aiiy ?o aiia?aoei?.
Iiaeaeuaiiy a ia/iioei iiaiia?iaeaeaieo oaa?ei eieueeinoi noaeei c
PNA+- aiaeioie?ioeeoaie iiaea aooe iciaeith eiiiainaoi?ii? ?aaeoei?
i?aaiicio, yea caaacia/o? aiaeooiooaaiiy iioiieiai/ii iaaeic?ieeo
e?ioi?aeieo eeioei aianeiaeie iayaiinoi ia iiaa?oii ye eiioioeeoia, oae
i aiaeioaeiioeeoia, iaeiieiaiieo ?aoeaioi?ia (?iaeinii I.A., Oiii?eiaa
I.A., O?ooaeei A.A., 1986). Ioaea, ciiia ?aoeaioi?iiai aia?aoo
aiaeioaeith iiaea aooe iaeiei c aieiaieo iaoaiiciia i?eieiaiiy
ooieoeiiioaaiiy ia/iiee ye oeaio?aeueiiai eiioi?aeiiai i?aaio.
Aioo?ioiuei/a?aaia aaaaeaiiy aioeaaiia i?enei?th? oeae i?ioean.
Iiaeiaii ?acoeueoaoe ciiie aeeiaiiee PNA-iiceoeaieo eiioioeeoia
aianeiaeie i?aiaoaeueii? aioeaaiii? noeioeyoei? niinoa?iaaee a oeioni
iiaiia?iaeaeaieo (Aieioei I.A., ?aaiia I.?.,1997). O aiaeiiaiaeue ia
aioo?ioiuei/a?aaia aioeaaiia iiae?aciaiiy c aeei/eiai? caeice
i?enei?thaaeanue aiia?aoeiy eiioioeeoia. *a?ac oa, ui a iaio i?aaiao
aeyaeythoueny iaeiioeiii ciiie o aeeiaiioei eaeoei-iiceoeaieo
eiioioeeoia, iiaeia c?iaeoe aeniiaie i?i inioaaiiy caeiiiii?ii? ?aaeoei?
oeaio?aeueieo i?aaiia iioii? nenoaie ia /oaea??aei? aioeaaie o
aioaiaoaeueiiio ia?iiaei.
Ia oiii i?eaii/aiiy eiioioeeoiiiaoe/ii? ooieoei? oa cieeiaiiy
eiioioeeoia, yei ? oaeoi?ii ii?oiaaiaco i?aaiia (Aaaaaaa.A.A., 1985), a
ia/iioei aiaeaoaa?oueny ooieoeiiiaeueia ia?aaoaeiaa oa aeoeaaoeiy iioeo
ooieoeie, yei ? i?eoaiaiieie ?e iiney ia?iaeaeaiiy, ui
noi?iaiaeaeo?oueny ciiiith oa?iiiia oi?ioaaiiy no?oeoo?ieo iaeeieoeue
i?aaio.
No?oeoo?ii-ooieoeiiiaeueii iaeieoei ia/iiee iiaiia?iaeaeaieo uo?ia ia
ia?oo aeiao aeeooy aoee ua ia noi?iiaaii. Ia/iieiai aaeee ia ae?aaeaii.
C ia?oi? aei /ioe?iaaeoeyoi? aeiae iinoiaoaeueiiai aeeooy ieiua
ia/iieiaeo /anoi/ie iinooiiai c?inoaea, c i?i?iooaaiiyi aei
oeaio?aeueieo aai ia/iieiaieo aaeie, yei neeaaeaeeny c aeaio-o?ueio
aaiaoioeeoia o aeiaaeeio. Oei no?iee aiaeiiaiaeathoue ia?oiio aoaio
noaiiaeaiiy no?oeoo?ii-ooieoeiiiaeueieo iaeeieoeue ia/iiee uo?ia
(Aaoaaee?ia O.I.,1998). C /ioe?iaaeoeyoi? aei o?eaeoeyoi? aeiae aeeooy o
iioaeoieo uo?ia niinoa?iaaee oii?yaeeoaaiiy ia/iieiaio aaeie, ?o /ioeo
i?i?ioaoeith oa c?inoaiiy na?aaeiuei? ieiui no?oeoo?ieo iaeeieoeue
ia/iiee. Ia o?eaeoeyoo aeiao o iioaeoieo oaa?ei aeyaeyeeny ia?oi
ia/iieiai /anoi/ee, ye? ca oi?iith iaaeeaeaeenue aei ainueieeooieeia,
ioi/aieo o?ueiia-/ioe?ia ia/iieiaeie o?iaaeaie, ui ? oa?aeoa?iith
iciaeith aoaeiae aei?ineeo oaa?ei.
Ia ia?oo aeiao iinoiaoaeueiiai aeeooy o uo?ia, yeei aaiaeeee
iioiiaeiaoeii oa aaeoeeio ia?ioeoo, na?aaeiy ieiua ia/iieiaeo /anoi/ie
(aiaeiiaiaeii 1.56o104 iei2 oa 1,83o104 iei2) aoea aieueoith, iiae o
iioaeoieo oaa?ei (1.41o104 iei2), ui aiaeia?aaea? i?enei?aiiy oi?ioaaiiy
ia?aioiie ia/iiee. Ii/eiath/e c o?aouei? aeiae aeeooy i a iiaeaeueoi
no?iee o uo?ia, yeei aaiaeeee iioiiaeiaoeii, ieiua /anoi/ie
c?iaith?oueny c iieacieeii iioaeoii? a?oie, oiaei ye o oaa?ei, yeei
aaiaeeee aaeoeeio ia?ioeoo, aiia i?iaeiaaeo? caeeoaoenue aieueo aenieith
oaeiae ia i'yoo oa nueiio aeiao. Ii/eiath/e c iaeeiaaeoeyoi? aeiae oa
aei eiioey ia?oiai iinyoey ?icaeoeo a iaio aenia?eiaioaeueieo a?oiao
aiaeii/a?oueny aieueo oii?yaeeiaaia, iiae o iioaeoieo, ?icoaooaaiiy
ia/iieiaeo aaeie, ui aiaeia?aaea? i?enei?aiiy oaiiia i?ioeano
oi?iiooai?aiiy /anoi/ie.
Aiaenooiinoue /anoi/eiai? aoaeiae ia/iiee iiaiia?iaeaeaieo uo?ia
iia'ycaii ic oa?aeoa?ii ?aei oaa?ei, yei aei eiioey ae?oaiai oeaeiy
aeeooy aaeeaathoue aeeeth/ii iaoa?einueea iieiei, a iioii ii/eiathoue
ia?aoiaeeoe ia naiinoieia oa?/oaaiiy, ui aiaeiiaiaea? e?oa?aoo?iei
aeaiei (Caiaaeithe E.I., 1983).
Oea i?ecaiaeeoue aei ?iceiai aeiiaioa?iiai iaaaioaaeaiiy ia ia/iieo.
Iacaeaaeii aiae a?oie oaa?ei, ciiia caniao oa?/oaaiiy oa i?ia?aneaia
ciaioaiiyi eiioi?aeiiai eiiiiiaioo i?aaio, ? aieiaieie /eiieeaie
ii?oiaaiaco ia/iiee.
O iiaiia?iaeaeaieo aenia?eiaioaeueieo uo?ia, ia oiii aieueo aeniei?,
iiae o iioaeoieo, aiaeiinii? iane i?aaio oa na?aaeiuei? ieiui iaeaooiio
ia/iieiaeo /anoi/ie, aiaeii/aeanue aiaeiiaiaeii aieueoa ieiua ia/iieiaeo
noaeei. O oaa?ei, yei iaea?aeoaaee aaia-aeiaoeii, niinoa?iaaeanue eeoa
oaiaeaioeiy aei ?? c?inoaiiy, a, ii/eiath/e c o?aouei? aeiae i iaaeaei,
aiia i?aeoe/ii c?iaith?oueny c eiio?ieueieie iieacieeaie. O uo?ia, yei
io?eioaaee aaeoeeio ai?ono ia?ioeoo, ia ia?oo aeiao aeeooy aiaeiinia
ieiua noaeei ia/iiee aoea aeinoiai?ii aeuith, iiae o iioaeoieo
(aiaeiiaiaeii 25.5+1.8% i 18.3+1.8%). Neioni?aee ?icoe?aii. Ia i`yoo oa
nueiio aeiao aeeooy aaee/eia aiaeiinii? ieiui noaeei a ia/iioei oeeo
uo?ia caeeoaeenue i?aeoe/ii ia iaeiiio ?iaii. Ii/eiath/e c eiioey
ae?oaiai oeaeiy aiia c?iaith?oueny ic iieacieeii eiio?ieth.
Iaeiei c aaaeeeaeo oaeoi?ia o i?ioeanao c?inoaiiy iane i?aaio oa
oi?ioaaiii eiai no?oeoo?ieo eiiiiiaioia ? noai i?ieioa?aoeaii?
aeoeaiinoi eeioei (Aiaaeaiiaa I.N.,1990), yeee caeaaeeoue aiae
niiaaiaeiioaiiy eeioeiii? oa aioo?ioiueieeioeiii? oi?i ?aaaia?aoei?
aaiaoioeeoia oa ia?ic?eaii iia'ycaiee ic iniaeeainoyie iaiiio
ioeea?iiaeo eeneio. O uo?ia iioaeoii? a?oie ia i?ioyci ia?oiai iinyoey
aeeooy ia aeyaeaii ai?iaiaeieo eieeaaiue o aeeiaiioei aiinoo AeIE a
yae?ao aaiaoioeeoia, ui aiaeiiaiaea? eioa?aoo?iei aeaiei (Iaeaaaa
A.O.,O?ieoeeay A.E., Aai?ia E.A., 1985). O uo?ia aenia?eiaioaeueieo a?oi
ii/eiath/e c ia?oi? aeiae aeeooy niinoa?iaaeinue oaiaeaioeiy aei
caieueoaiiy aiinoo AeIE a yae?ao (2.8+0.5 o.i. a ae?oaie a?oii oa
2.9+0.2 o.i. a o?aoiie a?oii) o ii?iaiyiii c iioaeoieie oaa?eiaie
(2,8+0.3 o.i.) Aei o?aouei? aeiae o uo?ia, yei io?eioaaee aaeoeeio
ia?ioeoo, oeae iieaciee ciaioeany, aea ia i`yoo oa nueiio aeiao ciiao
niinoa?iaaeanue oaiaeaioeiy aei eiai c?inoaiiy. C /ioe?iaaeoeyoi? aei
o?eaeoeyoi? aeiae ai?ia?aeieo aiaeiiiiinoae o aiinoi AeIE a yae?ao
aaiaoioeeoia iiae oaa?eiaie eiio?ieueieo oa aenia?eiaioaeueieo a?oi ia
niinoa?iaaee.
Ia oiii c?inoaiiy aiinoo AeIE a yae?ao o oaa?ei aenia?eiaioaeueieo a?oi
c ia?oi? aeiae aeeooy aiaeii/aee aieueo aenieee, iiae o iioaeoieo, aiino
iioioe/ii aeoeaieo aaiaoioeeoia (1.4+0.4% i?ioe 0,8+0.2%), oa iaioo
aiaeiinio eieueeinoue iieiioeeaa?ieo eeioei (2.0+0.9% i?ioe 4,3+0.4%) ia
i?ioyci ia?oiai oeaeiy. Aei o?eaeoeyoi? aeiae aeeooy oei aiaeiiiiinoi
iiaaethaaeenue. C?inoaiiy o iiaiia?iaeaeaieo uo?ia aenia?eiaioaeueieo
a?oi aiaeiinii? eieueeinoi iioioe/ii aeoeaieo aaiaoioeeoia ei?aeth? c
aiaeiiaiaeiei caieueoaiiyi aiinoo AeIE a yae?ao, iiaeaeuaiiy neioaco
yei? ia?aaeo? aeoeaaoei? i?ieioa?aoeaieo i?ioeania o eeioeiao, ui ?
iiaenoaaith
i?enei?aiiy oi?ioaaiiy ia?aioiie ia/iiee aianeiaeie aioo?ioiuei/a?aaii?
aei? aioeaaiia.
Iioaa?aeueiei iieacieeii aiaeia?aaeaiiy ooieoeiiiaeueiiai noaio i?aaio ?
aeieiaaineioaoe/ia ooieoeiy ia/iiee, yeo ioeiithaaee ia iniiai
?iciiaeieaiiy oa aeeiaiiee aeeth/aiue aeieiaaio a oeeoiieacii
aaiaoioeeoia.
I?yii? i?iii?oeiiiaeueii? caeaaeiinoi iiae iaeiie/aiiyi aeieii?ioa?aeia,
yei aeyaeyeeny ca aeiiiiiaith O?E-?aaeoei? oa ?aaeoei? Aanoa, ia
anoaiiaeaii. O iiaiia?iaeaeaieo oaa?ei aenia?eiaioaeueieo a?oi o
ii?iaiyiii c iioaeoieie niinoa?iaaee caieueoaiiy aiinoo Oeoo-iiceoeaiiai
iaoa?iaeo a iaia?aiao oa oeeoiieacii aaiaoioeeoia, niieo/iie oeaieii
eainoee e ii?oaeueieo o?aeoia. A aiaeioaei? noaeei aiino
aeieii?ioa?aeia, iaaiaee, aoa iaioei.
Ia ia?oo aeiao iiney ia?iaeaeaiiy o oeeoiieacii aaiaoioeeoia iioaeoieo
uo?ia niinoa?iaa?oueny ?iaiiii?iee ?iciiaeie aeieiaaio o aeaeyaei
e?aiaeue na?aaeiueiai ?icii?o (68.9% a?ae caaaeueii? e?eueeino?
aaiaoioeeo?a). Aei o?aouei? aeiae aeeooy aiaeii/aee iiyao aaiaoioeeoia,
iicaaaeaieo aeeth/aiue aeieiaaio (3.5+1.2%), ui aeaco? ia i?enei?aiiy
iaoaaieicio aeieiaaio. Aiino eeioei c aeeth/aiiyie o aeaeyae? ae??aieo:
na?aaei?o oa aaeeeeo a?aioe i?aeoe/ii ia ciiithaaany ia i?ioyci ia?oeo
aeaio oeaeiia aeeooy. Iiney /ioe?iaaeoeyoi? aeiae aeeooy a ia/iioei
iioaeoieo uo?ia aiaeii/aeanue noaaieicaoeiy niiaaiaeiioaiiy eeioei ic
aeeth/aiiyie aeieiaaio ?iciiai ?icii?o, ui na?ae/eoue i?i anoaiiaeaiiy
?iaiiaaae iiae i?ioeanaie aeieieico oa aetheiiaiaaiaco.
O iiaiia?iaeaeaieo uo?ia, yei io?eioaaee aioeaaie, ia aiaeiiio aiae
iioaeoieo uo?ia, a iaaeieiii?oaeueieo ciiao ia/iiee niinoa?iaaee aaeeeo
eieueeinoue aaiaoioeeoia, yei ia iaee aeeth/aiue aeieiaaio a oeeoiieacii
(aei 10.0%). ?o aiino i?iaeiaaeoaaa caeeoaoeny aieueo aenieei, iiae o
iioaeoieo oaa?ei, ia i?ioyci o?aouei?-nueiii? aeiae aeeooy. Eieueeinoue
aaiaoioeeoia c aeeth/aiiyie aaeeeiai ?icii?o, iaaiaee, aei i'yoi? aeiae
aeeth/ii aoea iaioith (5.0% i?ioe 11.9%), a iiney nueiii? - aieueoith,
iiae o eiio?iei. Ii/eiath/e c eiioey ae?oaiai oeaeiy iiney ia?iaeaeaiiy
o uo?ia iaio aenia?eiaioaeueieo a?oi niiaaiaeiioaiiy eeioei, ui iinoeee
aeeth/aiiy aeieiaaio ?iciiai ?icii?o, oa ?o ?icoaooaaiiy a iaaeao
ia/iieiaeo /anoi/ie i?aeoe/ii c?iaith?oueny c iieacieeaie iioaeoieo
uo?ia.
C?inoaiiy oaiiia ?iciaaeo aoaeaaiaeia o ia/iioei aenia?eiaioaeueieo
oaa?ei ? na?ae/aiiyi aieueo ?aiiueiai ii/aoeo ooieoeiiioaaiiy i?aaio a
?aiiueiio iinoiaoaeueiiio ia?iiaei iioiaaiaco. ?aiiy aeoeaaoeiy
aeieiaaiiiai iaoaaieicio a aaiaoioeeoao iiney aioo?ioiuei/a?aaii?
iioiicaoei? iiaea aooe aeiaeaoeiaei oaeoi?ii i?enei?aiiy oii?yaeeoaaiiy
/anoi/ie ia/iiee o aenia?eiaioaeueieo oaa?ei. Ciiia caniao oa?/oaaiiy c
ae?oaiai oeaeiy iiaaeth? aeyaeaii ciiie o ii?iaiyiii c iioaeoieie
uo?aie.
AENIIAEE.
1. Aioo?ioiuei/a?aaia aaaaeaiiy aioeaaiia,iacaeaaeii aiae ?o i?e?iaee,
aeeeeea? o?aiceoi?io aaiaoiiaaaeith iiaiia?iaeaeaieo, yea iiney
aaaaeaiiy aaia-aeiaoeiio o?eaa? aei o?aouei? aeiae aeeooy, a iiney
aaaaeaiiy aaeoeeie ia?ioeoo - aei i'yoi?.
2. Eiioi?aeiee eiiiiiaio ia/iiee iiaiia?iaeaeaieo uo?ia caeia? aei 18%
ieiui ia/iiee i i?aaenoaaeaiee iiiaeeiieeie eiioioeeoaie oa ?o
neoi/aiiyie aeaio aeaeia: iaeii neoi/aiiy eieaeiciaaii iaaeiei
ia/iieiaeo noaeei, iioi ?icoaoiaaii iiae aaiaoioeeoaie. ssaeua
eiioioeeoiiiaco a ia/iioei iioaeoieo uo?ia i?eieiythoueny i?ioyaii
ia?oeo aeaio oeaeiia aeeooy. Aioo?ioiuei/a?aaia aaaaeaiiy aioeaaiia
ieiaeai, iacaeaaeii aiae ?o aeaeo, aeeeeea? ciaioaiiy aiinoo eiioi?aeii?
oaeeie o iiaiia?iaeaeaieo oa iiaeaeueoa i?eaii/aiiy eiioioeeoiiiaoe/ii?
ooieoei? ia/iiee oeyoii i?enei?aiiy aiia?aoei? c i?aaio eiioioeeoia.
3. ?aoeaioi?e aei eaeoeio a?aoino a ia/iioei o iiaiia?iaeaeaieo uo?ia
aeyaeythoueny ia iaia?aii oa a oeeoiieacii aiaeioaeiioeeoia,
eiioioeeoia, aiioaeiioeeoia aeia/ieo i?ioie oa ci?eiiiaeiaieo
?aoeeoeiaiaeioaeiioeeoia. C ia?oi? ii o?aoth aeiao aeeooy ii/eia?oueny
c?inoaiiy aiinoo PNA+-aiaeioaeiioeeoia c iaeii/aniei ciaioaiiyi
PNA+-eiioioeeoia, yei iiaiinoth cieeathoue aei /ioe?iaaeoeyoi? aeiae.
Aioo?ioiuei/a?aaia aaaaeaiiy aioeaaiia aeeeeea? ia ia?oo aeiao aeeooy
c?inoaiiy, ii?iaiyii c iioaeoieie, aiinoo PNA+-aiaeioaeiioeeoia oa
cieaeaiiy eieueeinoi PNA+-eiioioeeoia a ia/iioei. Iaaeaei oaiie
ciaioaiiy aiaeiinii? eieueeinoi PNA+-eiioioeeoia aeia?aaeaeathoue
iieacieee eiio?ieth.
4. O oi?ioaaiii ia/iieiaeo /anoi/ie iioaeoieo uo?ia ia?oiai iinyoey
aeeooy niinoa?iaa?oueny aeai noaaei?: ia?oa i?ioiea? ia oiii caanaiiy
aaiioeeoiiiaco oa eiioioeeoiiiaco a ia/iioei oa o?eaa? aiae ia?iaeaeaiiy
aei eiioey ae?oaiai oeaeiy aeeooy, oa?aeoa?ecoth/enue oi?ioaaiiyi
ia/iieiaeo aaeie; ae?oaa ii/eia?oueny iiney 14-? aeiae aeeooy oa
oa?aeoa?eco?oueny oii?yaeeoaaiiyi aaeie oa inoaoi/iei oi?ioaaiiyi
ia/iieiaeo /anoi/ie, yei aei eiioey ia?oiai iinyoey aeeooy i?eiathoue
oi?io ainueieeooieea, ioi/aiiai ia/iieiaeie o?iaaeaie. O oaa?ei, yeei
aaiaeeee aioeaaie, i?enei?th?oueny caaa?oaiiy ia?oi? noaaei?, ui
i?iyaey?oueny o a?eueo oaeaeeiio oi?ioaaiii ia/iieiaeo aaeie oa
caieueoaiiyi ieiui /anoi/ie.
5. Aioo?ioiuei/a?aaia aaaaeaiiy aioeaaiia iineeth? i?ieioa?aoeaio
aeoeaiinoue aaiaoioeeoia iiaiia?iaeaeaieo, ui noi?iaiaeaeo?oueny
c?inoaiiyi neioaco AeIE a yae?ao aei i'yoi? aeiae aeeooy, iaeii/aniei
c?inoaiiyi eieueeinoi iioicia aei o?aouei? aeiae oa ciaioaiiyi aiinoo
iieiioeeaa?ieo aaiaoioeeoia aei /ioe?iaaeoeyoi? aeiae aeeooy.
6. Iiae aieeaii aioo?ioiuei/a?aaiiai aaaaeaiiy aioeaaiia o uo?ia ia?oi?
aeiae aeeooy c'yaeythoueny aaiaoioeeoe, iicaaaeaii
aeeth/aiue
aeieiaaio, yei o iioaeoieo aeyaeythoueny c o?aouei? aeiae. A
iiaeaeueoiio niinoa?iaa?oueny i?enei?aia, o ii?iaiyiii c iioaeoieie,
iaeiie/aiiy aeieiaaio a aaiaoioeeoao aei 30-? aeiae.
NIENIE I?AOeUe, IIOAE?EIAAIEO CA OAIITH AeENA?OAOe??.
1. ?ieue aioo?eoo?iaiie aioeaaiiie noeioeyoeee a oi?ie?iaaiee
aenoea?iiaaaeee i?aaiia o iiai?iaeaeaiiuo aeeaioiuo /Aieioei I.A.,
Eaaiia I.A., Ua?aaeia I.N., A.A.Iaaeaaaeaa, Eou I.A. // Ii?oieiaey (A?o.
aiao.),- 1998 - N 3 - N. 32-33.
2. Ua?aaeia I.N., Aieioei I.A., Eaaiia I.A. Iniaaiiinoe iaeiieaiey
aeeeiaaia a aaiaoioeeoao iiai?iaeaeaiiuo e?un iinea aioo?eoo?iaiiai
aaaaeaiey aioeaaia // Ainiee ii?oieiai?, 1998 - O.4 - N 2 - N. 200-201.
3. Iniaeeainoi ii?oiaaiaco iaaeie?ieeiaeo caeic oa ia/iiee uo?ia o
?aiiueiio iinoiaoaeueiiio ia?iaei /Aieioei I.A., Ua?aaeia I.N., Aia/aiei
I.A., Eou I.A. // Cai?iee iaoe. noaoae “Aeooaeueii ieoaiiy
oa?iaoeaaoe/ii? oa iaaee/ii? iaoee oa i?aeoeee”, –Caii?iaeaey, 1997. –N.
207-211
4. Aieioei I.A., Ua?aaeia I.N. Iniaaiiinoe oi?ie?iaaiey ia/aie o
iiai?iaeaeaiiuo e?un iinea aioo?eoo?iaiiai aaaaeaiey aioeaaia // Ainiee
Aieioea?eianueeiai aea?aeaaiiai aa?a?iiai oiiaa?neoaoo. Iaoeiai noaooi
iiaeia?iaeii? iaoeiai-ae?iaie/i? eiioa?aioei? "Ii?oieiaiy - i?aeoe/iie
aaoa?eia?i? oa iaaeeoeeii", - 1998, - aei. 6. - *.1.- N.133-136.
5. Ua?aaeia I.N., Aia/aiei I.A. Iniaaiiinoe oi?ie?iaaiey eeioieaeiie
nenoaiu ia?aioeiaoiciuo i?aaiia e?un a ?aiiai iinoiaoaeueiii ia?eiaea //
Oaoiaa aeaeaiiy iaoeiaeo i?aoeue iaoa?iaeia ?? iaoeiiiaeueiiai eiia?ano
aiaoiiia, ainoieiaia, aia?iieiaia i oiiia?aoiaiaoiiia Oe?a?ie
"Aeooaeueii ieoaiiy ii?oieiai?" - Eoaainuee: AAO "EIAe", 1998, -
N.262-263.
6. Aieioei I.A. Ua?aaeia I.N., Iiainaeiaa I.A. Iniaaiiinoe no?iaiey
ia/aie e?un a ?aiiai iinoiaoaeueiii ia?eiaea iiiaaiaca eae
oeaio?aeueiiai eeioieaeiiai i?aaia // Iaoeiai aeineiaeaeaiiy c ieoaiue
iaaee/ii? aiieiai?, oiii?, aa?a?i? oa no/anieo oaoiieiai?e iaa/aiue.
Ui?i/iee. - Oa?iiiieue, 1997. - aeione 1, /ano. 1, –N.125-126.
7. Iniaaiiinoe ii?oiaaiaca i?aaiia a caaeneiinoe io iooae i?iieeiiaaiey
aioeaaiia a i?aaieci ieiaea /Ea?cia I.A., Ua?aaeia I.N.,Iaaeaaaeaa A.A.,
Iii?aaei I.E. // Nai?iee noaoae “Aeooaeueiua aii?inu iaaeeoeeiu e
aeieiaee”., – Aeiai?iiao?iane, 1996. – N.135 .
8. Eeioioeeou eae oaeoi? ii?oiaaiaca i?aaiia Aieioei I.A., Eaaiia I.A.,
Iiainaeiaa I.A., Ua?aaeia I.N.,Aia/aiei I.A., Iaaeaaaeaa A.A., Iii?aaei
I.E. // Iaoa?iaee iaoe. eiio. “Aeooaeueii ieoaiiy ii?oiaaiaco”. -
*a?iiaoei, 1996,- N. 76-77.
9. Ua?aaeia I.N.,Iiainaeiaa I.A., Aia/aiei I.A. Iniaaiiinoe ii?oiaaiaca
ia/aie, iaaeii/a/ieeia e naeacaiee e?un a ?aiiai iinoiaoaeueiii ia?eiae
iioiaaiaca //Iaoa?iaee iiaeia?iaeii? eiio. “Aeooaeueii ieoaiiy
ii?oieiai?”. - Oa?iiiieue, 1996,- N. 475 -477 .
10. I?eioeeiu neiiao?ee e aneiiao?ee a no?iaiee i?aaiia eeioieaeiie
nenoaiu. /Aieioei I.A., A?otho A.A., Eaaiia I.A., Iiainaeiaa I.A.,
Ea?cia I.A., Ia?ane/ A.I., Iii?aaei I.E. //I?eioeeie i?iii?oei?,
neiao?i?, no?oeoo?ii? aa?iiii? oa iaoaiaoe/iiai iiaeaethaaiiy a
ii?oieiai?. - Aiiieoey, 1997. - N.38.
11. Eco/aiea aeeyiey i?aia?aoa "A" ia eiioiioth nenoaio i?aaiecia a i?a-
e iinoiaoaeueiii ia?eiaea ?acaeoey. /Aieioei I.A., Iaco? E.A., Eiuo
A.A., Iaeuoaa A.A., A?aaeieeia A.I., Oieiaeiay A.E., Iiainaeiaa I.A.,
Eaaiia I.A. Ua?aaeia I.N., // Oacenu aeiee. iao/ii-i?aeo. eiio.
“Eaea?noaaiiua n?aaenoaa Oe?aeiu, neioac, iao/iua enneaaeiaaiey,
i?iecaiaenoai, ?aaeecaoeey”. - Oa?ueeia, 1992. - N. 186.
12. Aioo?eoo?iaiia aioeaaiiia aicaeaenoaea - oaeoi? ?acaeoey
aenoea?iiaaaeee iinea ?iaeaeaiey. /Aieioei I.A., Ea?cia I.A., Eaaiia
I.A., Ua?aaeia I.N.,Eou I.A., Iaaeaaaeaa A.A., Iiainaeiaa I.A.,
Ai?iaeie/aaa A.A. //Cai?ea oac ? iaoeiiiaeueiiai eiia?ano Oe?aiie c
iioiieiaii, aea?aieiaii oa iioii?aaaieioaoeii. - Aeoooa, E?ei, 1998.- N.
42-43 .
AIIOAOeIss
Ua?aaeia I.N. Iniaeeainoi aoaeiae ia/iiee uo?ia a ?aiiueiio
iinoiaoaeueiiio ia?iiaei a ii?ii oa iiney aioo?ioiuei/a?aaiiai aaaaeaiiy
aioeaaiia (aiaoiii-aenia?eiaioaeueia aeineiaeaeaiiy). - ?oeiien.
Aeena?oaoeiy ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa iaaee/ieo iaoe
ca niaoeiaeueiinoth 14.03.01 - ii?iaeueia aiaoiiiy. - Oa?iiiieuenueea
aea?aeaaia iaaee/ia aeaaeaiiy ii. ?.ss.Ai?aa/aanueeiai, Oa?iiiieue,
1999.
?iaioa i?enay/aia aeineiaeaeaiith oi?ioaaiiy ia/iiee uo?ia oa ??
eiioi?aeiiai eiiiiiaioo ia i?ioyci ia?oiai iinyoey iinoiaoaeueiiai
aeeooy ia oiii i?eaii/aiiy ?? ooieoei? ye eiioi?aeiiai i?aaio a ii?ii oa
iiney aioo?ioiuei/a?aaiiai aaaaeaiiy aioeaaiia ia 18-o aeiao
aioaiaoaeueiiai ?icaeoeo.
Anoaiiaeaii, ui aioo?ioiuei/a?aaia aaaaeaiiy aioeaaiia aeeeeea? o
iiaiia?iaeaeaieo o?aiceoi?io aaiaoiiaaaeith oa ciiith? no?iee oi?ioaaiiy
no?oeoo?ii-ooieoeiiiaeueieo iaeeieoeue ia/iiee, ui noi?iaiaeaeo?oueny
i?enei?aiiyi aiia?aoei? c i?aaio eiioioeeoia.
Eeth/ia? neiaa: ia/?iea, e?ioi?aeia oeaieia ia/?iee, aioo??oiuei/a?aaia
aioeaaiia noeioeyoe?y.
AIIIOAOeEss
Ua?aaeia I.N. Iniaaiiinoe ii?oiaaiaca ia/aie e?un a ?aiiai
iinoiaoaeueiii ia?eiaea a ii?ia e iinea aioo?eoo?iaiiai aaaaeaiey
aioeaaiia (aiaoiii-yenia?eiaioaeueiia enneaaeiaaiea). - ?oeiienue.
Aeenna?oaoeey ia nieneaiea iao/iie noaiaie eaiaeeaeaoa iaaeeoeeineeo
iaoe ii niaoeeaeueiinoe 14.03.01 - ii?iaeueiay aiaoiiey. -
Oa?iiiieuenueeay ainoaea?noaaiiay iaaeeoeeineay aeaaeaiey ei.
E.ss.Ai?aa/aaneiai, Oa?iiiieue, 1999.
Iaoa?eaeii enneaaeiaaiey iineoaeeea ia/aiue 187 eioaeoiuo e
yenia?eiaioaeueiuo e?un ia?aiai ianyoea iinoiaoaeueiie aeecie. A
yenia?eiaioa eniieueciaaee iiaeaeue aioo?eoo?iaiiai aaaaeaiey aioeaaiia
ia 18-a nooee ieiaeiiai ia?eiaea ?acaeoey. A ea/anoaa aioeaaiia auee
aua?aiu eiioiiaeiaoeei /aeiaaea e nooay aeeaay aaeoeeia ae?oniiai
ia?ioeoa. Eiio?ieueiay a?oiia aeeaioiuo iieo/aea oeceieiae/aneee
?anoai?.
Ioiineoaeueiay ianna ia/aie eioaeoiuo e?un ia i?ioyaeaiee ia?auo aeaoo
iaaeaeue aeecie nieaeaeanue n iineaaeothuei iiauoaieai aei o?eaeoeaouo
nooie. O iiai?iaeaeaiiuo e?un iaaeo yenia?eiaioaeueiuo a?oii ia ia?aua
nooee aeecie ioia/aee o?aiceoi?ioth aaiaoiiaaaeeth, aeeeoaeueiinoue
eioi?ie caaenaea io aeaea aaiaeeiiai aioeaaia.
Eeioieaeiue eiiiiiaio ia/aie iiai?iaeaeaiiuo e?un i?aaenoaaeai
aaeeie/iuie eeioioeeoaie e eeioieaeiuie neiieaieyie, ioiineoaeueiay
ieiuaaeue eioi?uo i?ia?anneaii nieaeaeanue aei eiioea aoi?ie iaaeaee. Ia
ia?aua nooee aeecie a ia/aie yenia?eiaioaeueiuo aeeaioiuo ioia/aee
iaiueooth, /ai o eioaeoiuo, ieiuaaeue eeioieaeiie oeaie. Oaiiu aunaeaiey
eeioioeeoia ec i?aaia ia iaeie-aeaia nooie iia?aaeaee iieacaoaee
eiio?iey, e yeeieiaoeey caaa?oaeanue e 11-i nooeai aeecie.
?aoeaioi?u e eaeoeio a?aoena a aeaea eiioeaauo inoaoeia -D-aaeaeoicu,
ii?aaeaeyeenue, i?aeiouanoaaiii, a eeioioeeoao e yiaeioaeeioeeoao, a
oaeaea, a iaiueoai eiee/anoaa, a eeaoeao Eoioa?a e yieoaeeioeeoao
aeae/iuo i?ioieia. PNA+-eeioioeeou - eiioiieiae/anee iac?aeua. O
eioaeoiuo aeeaioiuo ioia/aee ia?aoioth i?iii?oeeiiaeueioth caaeneiinoue
iaaeaeo eiee/anoaii PNA+-eeioioeeoia, eiee/anoai eioi?uo iinoaiaiii
oiaiueoaeinue, e PNA+-yiaeioaeeioeeoia. O aeeaioiuo, eioi?ui aaiaeeee
aioeaaiu, ia ia?aua nooee aeecie iaaethaeaee cia/eoaeueii iaiueoaa, /ai
o eioaeoiuo, /enei PNA+-eeioioeeoia, e aieueoaa - PNA+-yiaeioaeeioeeoia.
E 14-i nooeai aeecie, eiaaea caeai/eaaaony yiea?aoeey eeioioeeoia ec
ia/aie eioaeoiuo e?un, PNA+-eeioioeeou ia auyaeyeenue.
Ia oiia nieaeaiey ieiuaaee eeioieaeiie oeaie ia/aie iaaethaeaeinue
oi?ie?iaaiea no?oeoo?ii-ooieoeeiiaeueiuo aaeeieoe i?aaia, /oi
nii?iaiaeaeaeinue oii?yaei/aieai ?aniieiaeaiey ia/aii/iuo aaeie e
oaaee/aieai n?aaeiae ieiuaaee ia/aii/iuo aeieae. O e?un
yenia?eiaioaeueiuo a?oii ia ia?aua nooee aeecie ioia/aee aieueooth, /ai
o eioaeoiuo aeeaioiuo, ieiuaaeue aeieae, oi?ie?iaaiea eioi?uo
inouanoaeyeinue aieaa auno?uie oaiiaie, /ai o eioaeoiuo. O
iiai?iaeaeaiiu e?un, iieo/aaoeo aioeaaiu, iaaethaeaee oaaee/aiea
niaea?aeaiey AeIE a yae?ao aaiaoioeeoia, /oi nii?iaiaeaeaeinue
iiauoaieai ioiineoaeueiiai /enea aaiaoioeeoia n oeao?aie ieoica e
oiaiueoaieai eiee/anoaa iieeieieaeiuo eeaoie ii n?aaiaieth n eiio?ieai.
Auoaoeacaiiua eciaiaiey nii?iaiaeaeaeenue aieaa ?aiiae aeoeaaoeeae
aeeeiaaineioaoe/aneie ooieoeee ia/aie, /oi i?iyaeyeinue a iiauoaiee
?aniaaea aeeeiaaia a aaiaoioeeoao. Aieueoeinoai eciaiaiee
ieaaee?iaaeenue iinea aoi?ie iaaeaee iinoiaoaeueiiai ?acaeoey eee e
eiioeo ia?aiai ianyoea aeecie - n?iea ieii/aoaeueiiai caaa?oaiey
iea?aoeee ec ia/aie eeioieaeiie oeaie e ia?aoiaea aeeaioiuo e
naiinoiyoaeueiiio niiniao ieoaiey.
Eeth/aaua neiaa: ia/aiue, eeioieaeiay oeaiue ia/aie, aioo?eoo?iaiay
aioeaaiiay noeioeyoeey.
SUMMARY
Maxim S. Shcherbakov Peculiarites of rat`s liver morphogenesis within
early postnatal period normally and after intrauterine injection of
antigens (anatomical-experimental study). - Manuscript.
Thesis for scientific degree of Candidate of Medical Sciences.
Speciality: 14.03.01 - Normal Anatomy. - Ternopil State Medical Academy
named after I. Y.Gorbachevsky. - Ternopil, 1999.
This whork is dedicated to investigation of rat`s liver formation during
the first month of postnatal life on a background of its functioning as
lymphoid organ is being oppressed normally and after intrauterine
antigen injection on the 18th day of antenatal development.
It`s established that intrauterine injection of antigens caused
hepatomegalia of new-borns and changed terms of liver
structure-functional units formation that was accompanied by
acceleration of lymphocytes emigration from the organ.
Key-words: liver, lymphoid tissue of liver, intrauterine antigen
injection.
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